Preheat oven to 400°F. Bring large pot of water to a boil for the pasta.
Place tuna in a bowl and add eggs, breadcrumbs, pine nuts, Parm, parsley, oregano, lemon zest, salt, pepper, grated or pasted garlic and a fat drizzle of olive oil. Roll balls the size of golf balls and place them on a parchment-lined baking sheet. Bake to golden brown and hot through, 15-18 minutes. Remove from oven.
Meanwhile, heat 2 tablespoons olive oil in large skillet over medium heat and sauté onions and sliced or chopped garlic for 5 minutes. Add wine, crushed tomatoes and passata or tomato purée. Season with salt, add torn basil and simmer 20 minutes. Add balls to sauce when they come out of the oven.
Cook pasta to al dente in salted water. Reserve 1 cup of starchy, salty water before draining.
Remove meatballs from sauce to a serving dish. Toss pasta with sauce, a handful of Parm, a drizzle of EVOO and the starchy, salty water to coat. Transfer pasta to serving bowl and pass with tuna balls and extra cheese alongside.